Alois Čelechovský1 &
Jaroslav
Bosák2
|
1 Department
of Zoology and Laboratory of Ornithology, Faculty of Science, Palacký
University, tř. Svobody 26, CZ-771 46 Olomouc, Czech Republic;
celechov@prfnw.upol.cz
2 Ecological
Consulting, a. s., Na Střelnici 48, CZ-779 00 Olomouc, Czech Republic;
jaroslav.bosak@ecological.cz
|
The family Bombyliidae includes several
significantly differentiated subfamilies. Many different opinions concerning
their taxonomic status and position have been formulated in last few decades
(e.g.
Greathead & Evenhuis 1997,
Zaitzev 1991). In the PCV2 (Bosák
1997a,
b), the Phthiriidae was ranked as
a separate family, however, here we are again classifying the Phthiriinae as
a subfamily of the family Bombyliidae (c. f.
Bosák 2006), in agreement
with the overall approach to the family within the framework of the
Fauna
Europaea (Greathead 2007).
Body size of adults ranging from 0.9 to 27.0 mm. Body in most
cases covered with hairs, bristles or pollinosity. Wings absolutely hyaline
or with diverse colour patterns frequently covering the entire wing surface.
Head rounded and convex anteriorly; occiput flat or tumid with a single
occipital foramen or concave with occipital foramen divided and with a deep
groove behind the ocellar triangle. Ocelli present. Proboscis short (Anthracinae,
Lomatiinae) to long, as much as four times as long as head (e.g. Bombyliinae,
Phthiriinae). Legs slender and long, empodium absent, and pulvilli present or
absent. Thorax uniform in structure. Abdomen wide, rounded, cylindrical or
short to elongate-conical. The adult morphology is treated in detail in a
number of papers (e.g.
Greathead
& Evenhuis 1997,
Hull 1973). The adults are heliophilous and suck nectar from flowers. The
larvae are parasites or hyperparasites, and their hosts are various insect
species from the orders Coleoptera, Diptera, Hymenoptera, Lepidoptera,
Neuroptera and Orthoptera.
The total number of species worldwide is
approximately 4000. Altogether 335 species of the family are currently known
from Europe (Greathead 2007). The preset
checklist consists of 55 species from the Czech Republic and Slovakia: 51
from the Czech Republic (43 from Bohemia, 43 from Moravia), and 47 from
Slovakia (Bosák
1997a,
b,
2006,
Čelechovský 2007, Greathead
et al. 2005). The species-richness of the bee-fly fauna is well known in
both the Czech Republic and Slovakia.
|
References
[1] Bosák J. 1997a: Phthiriidae.
In Chvála M. (ed.): Check List of Diptera (Insecta) of the Czech and
Slovak Republics. Karolinum – Charles University press, Praha, pp. 45-46.
[2] Bosák J. 1997b:
Bombyliidae. In Chvála M. (ed.): Check List of
Diptera (Insecta) of the Czech and Slovak Republics. Karolinum – Charles
University press, Praha, pp. 45.
[3]
Bosák J. 2006: Bombyliidae Latreile 1802. In Jedlička L., Stloukalová V
& Kúdela M. (eds): Chcecklist of the Czech Republic
and Slovakia. Electronic version 1. <http://zoology.fns.uniba.sk/diptera> +
CD-ROM: ISBN 80-969629-0-6. Retrieved
04.08.2009.
[4]
Čelechovský A. 2007: Faunistic records from Czech Republic and Slovakia.
Bombyliidae. In Stloukalová V. (ed.): Dipterologica bohemoslovaca Vo. 14, Acta Zoologica Universitatis Comenianae 47: 254.
[5] Greathead D. 1995: A
review of the genus Bombylius Linnaeus sensu lato (Diptera: Bombyliidae) from
Africa and Eurasia. Entomologica scandinavica 26: 47-66.
[6]
Greathead D. 2007:
Fauna Europaea: Bombyliidae. In Pape T. (ed.): Fauna Europaea: Diptera,
Brachycera. Fauna Europaea version 1.3.
<http://www.faunaeur.org>.
Retrieved 04.08.2009.
[7] Greathead D.J. & Evenhuis
N.L. 1997: Bombyliidae. In Papp L. & Darvas B. (eds): Contributions
to a Manual of Palaearctic Diptera (with special reference to flies of
economic importance).
Vol. 2. Nematocera and Lower Brachycera. Science Herald, Budapest,
487-512 pp.
[8]
Greathead D., Gregor T., Barták M., Kubík Š. & Bosák J. 2005: Bombyliidae. In
Barták M. & Kubík Š. (eds):
Diptera of
Podyjí National Park and its Environs.
Česká zemedělská univerzita v Praze, 137-140 pp.
[9] Hull F.M. 1973:
Bee flies of the World. The genera of the family Bombyliidae.
Bulletin of the United States National Museum
286: 1-687.
[10] Zaitzev V.F. 1991: The
phylogeny and system of Diptera of the superfamily Bombylioidea (Diptera).
Entomological Review 71(4): 94-114.
|
ANTHRACINAE |
|
|
|
|
|
|
|
|
|
|
|
|
|
Anthrax Scopoli, 1763 |
|
|
|
|
|
|
|
|
|
|
|
|
anthrax
(Schrank, 1781) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
binotatus
Wiedemann in Meigen, 1820 |
CZ |
(B |
M |
) |
|
|
|
|
|
|
|
|
trifasciatus
Meigen, 1804 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
varius
Fabricius, 1794 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
Spogostylum Macquart, 1840 |
|
|
|
|
|
|
|
|
|
|
|
|
aethiops
(Fabricius, 1781) |
CZ |
( |
M |
) |
SK |
|
BOMBYLIINAE |
|
|
|
|
|
|
|
|
|
|
|
|
|
Anastoechus Osten-Sacken, 1877 |
|
|
|
|
|
|
|
|
|
|
|
|
nitidulus
(Fabricius, 1794) |
CZ |
(B |
|
) |
SK |
|
|
|
|
|
Bombylella Greathead, 1995 |
|
|
|
|
|
|
|
|
|
|
|
|
atra (Scopoli,
1763) |
CZ |
(B |
M |
) |
SK |
in PCV2 in genus Bombylius Linnaeus, 1758 [5] |
|
|
|
|
Bombylisoma Rondani, 1856 |
|
|
|
|
|
|
|
|
|
|
|
|
minimum
(Scopoli, 1772) |
|
|
|
|
SK |
|
|
|
|
|
|
|
nigriceps (Loew,
1862) |
|
|
|
|
SK |
|
|
|
|
|
Bombylius Linnaeus, 1758 |
|
|
|
|
|
|
|
|
|
|
|
|
analis Pôivier, 1798 |
CZ |
(B |
M |
) |
SK |
in PCV2 and ECV1 as undatus Mikan, 1796 [6] |
|
|
|
|
|
|
canescens Mikan, 1796 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
cinerascens
Mikan, 1796 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
discolor Mikan,
1796 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
fimbriatus
Meigen, 1820 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
fulvescens
Wiedemann in Meigen, 1820 |
CZ |
( |
M |
) |
SK |
|
|
|
|
|
|
|
major Linnaeus,
1758 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
medius Linnaeus,
1758 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
minor Linnaeus,
1758 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
nubilus Mikan,
1796 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
quadrifarius
Loew, 1855 |
CZ |
( |
M |
) |
SK |
|
|
|
|
|
|
|
undatus Mikan,
1796 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
posticus
Fabricius, 1805 |
CZ |
(B |
M |
) |
SK |
in PCV2 and ECV1 as B. vulpinus Wiedemann in Meigen, 1820 [6] |
|
|
|
|
|
|
venosus Mikan,
1796 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
Conophorus Meigen, 1803 |
|
|
|
|
|
|
|
|
|
|
|
|
virescens
(Fabricius, 1787) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
Systoechus Loew, 1855 |
|
|
|
|
|
|
|
|
|
|
|
|
ctenopterus
(Mikan, 1796) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
Triplasius Loew, 1855 |
|
|
|
|
|
|
|
|
|
|
|
|
pictus (Panzer,
1794) |
CZ |
(B |
M |
) |
SK |
in PCV2 and ECV1 in genus Bombylius Linnaeus, 1758 [6] |
EXOPROSOPINAE |
|
|
|
|
|
|
|
|
|
|
|
|
|
Exhyalanthrax Becker, 1916 |
|
|
|
|
|
|
|
|
|
|
|
|
afer (Fabricius,
1794) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
Exoprosopa Macquart, 1840 |
|
|
|
|
|
|
|
|
|
|
|
|
capucina
(Fabricius, 1781) |
CZ |
(B |
|
) |
SK |
|
|
|
|
|
|
|
cleomene Egger,
1859 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
jacchus
(Fabricius, 1805) |
CZ |
(B |
|
) |
SK |
|
|
|
|
|
|
|
minos (Meigen,
1804) |
CZ |
( |
M |
) |
SK |
|
|
|
|
|
Hemipenthes Loew, 1869 |
|
|
|
|
|
|
|
|
|
|
|
|
maura (Linnaeus,
1758) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
morio (Linnaeus,
1758) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
velutinus
(Meigen, 1820) |
CZ |
(B |
|
) |
SK |
|
|
|
|
|
Thyridanthrax Osten-Sacken, 1886 |
|
|
|
|
|
|
|
|
|
|
|
|
fenestratus
(Fallén, 1814) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
Villa Lioy, 1864 |
|
|
|
|
|
|
|
|
|
|
|
|
cana (Meigen,
1804) |
CZ |
(B |
M |
) |
|
in PCV2 also as V. quinquefasciata (Wiedemann in Meigen, 1820) [6] |
|
|
|
|
|
|
cingulata
(Meigen, 1804) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
claripennis
(Kowarz, 1867) |
|
|
|
|
SK |
|
|
|
|
|
|
|
deludens
François, 1966 |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
fasciata
(Meigen, 1804) |
CZ |
(B |
|
) |
SK |
in PCV2 and ECV1 as V. circumdata (Meigen, 1820) [6] |
|
|
|
|
|
|
halteralis
(Kowarz, 1883) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
hottentotta
(Linnaeus, 1758) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
humilis (Ruthe,
1831) |
CZ |
(B |
|
) |
SK |
|
|
|
|
|
|
|
ixion
(Fabricius, 1794) |
CZ |
(B |
|
) |
SK |
|
|
|
|
|
|
|
modesta (Meigen,
1820) |
CZ |
(B |
M |
) |
|
|
|
|
|
|
|
|
occulta
(Wiedemann in Meigen, 1820) |
CZ |
(B |
|
) |
|
|
|
|
|
|
|
|
paniscus (Rossi,
1790) |
CZ |
(B |
M |
) |
SK |
|
LOMATIINAE |
|
|
|
|
|
|
|
|
|
|
|
|
|
Lomatia Meigen, 1822 |
|
|
|
|
|
|
|
|
|
|
|
|
erinnys Loew,
1869 |
|
|
|
|
SK |
|
|
|
|
|
|
|
lachesis Egger,
1859 |
CZ |
( |
M |
) |
|
|
|
|
|
|
|
|
lateralis
(Meigen, 1820) |
CZ |
(B |
M |
) |
SK |
|
|
|
|
|
|
|
sabaea
(Fabricius, 1781) |
CZ |
( |
M |
) |
SK |
|
PHTHIRIINAE |
|
|
|
|
|
|
|
|
|
|
|
|
|
Phthiria Meigen, 1803 |
|
|
|
|
|
|
|
|
|
|
|
|
canescens Loew,
1846 |
CZ |
( |
M |
) |
|
|
|
|
|
|
|
|
gaedei Wiedemann
in Meigen, 1820 |
CZ |
( |
M |
) |
|
|
|
|
|
|
|
|
minuta
(Fabricius, 1805) |
CZ |
(B |
M |
) |
|
|
|
|
|
|
|
|
pulicaria
(Mikan, 1796) |
CZ |
(B |
M |
) |
SK |
|
|
|