The Scatopsidae are a medium-sized
but still inadequately known family of derived nematocerous Diptera.
Adults are minute to small midges ranging from 0.5 to 4.0 mm in
length, generally dark in colour, sometimes more or less marked with
yellow. Body rather strongly
built, often quite elongated, with parallel-sided abdomen. Eyes
holoptic, forming a characteristic supraantennal bridge. Antenna
comparatively short, composed of two basal segments and a flagellum
of 5-10 flagellomeres that are generally wider than long.
Wing venation reduced, generally only anterior veins
pigmented. Larvae flattened, with a complete cephalic capsule,
peripneustic. All known larvae are saprophagous, living in a wide
variety of media (rotten plants and fruits, rotten wood, under
decaying bark of dead trees, soil, leaf litter, fungi, dung, etc) (Haenni
& Vaillant 1994).
The adults may be found in nearly all kinds of biotopes, but more
frequently in open, fresh and marshy areas.
The world fauna of Scatopsidae
comprises some 350 species (Haenni
1997), of which 102 species in 22 genera are known in Europe (Haenni
2007 and recent additions). 52 species are included in the present checklist: 46 from
the Czech Republic (41 from Bohemia and 42 from Moravia) and 43 from
Slovakia. Since the preceding PCV2 (Martinovský
1997b), which was mainly based upon the important papers by Martinovský (1997a)
for the Czech Republic and by Kozánek and Roller (1997)
for Slovakia, the number of species had increased by eight in the Czech
Republic (ten in Bohemia and seven in Moravia) while it had remained the
same in Slovakia. The rather numerous additional records may be
found in a set of recent faunistic papers (Haenni
& Barták
2000,
2006,
Haenni et al. 2005,
Martinovský 1998). On the other hand,
two nomenclatorial changes
have been proposed by Haenni (1998,
2002).
By contrast only one new record for Slovakia has been added (Haenni 2009)
since the previous electronic version (ECV1,
Haenni 2006).
The species richness of the scatopsid fauna is rather well known in
the Czech Republic and in Slovakia, but the number of species will
continue to grow as there are several new species awaiting
description.
The male and
female genital characters enable safe identifications to be made. A
key to the Palaearctic genera (both larvae and adults) may be found
in Haenni (1997),
where references to papers for identification to species level are
also included. The classification used here follows Amorim (1994)
and Chandler (1998),
while the nomenclature follows Haenni (2007).
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References
[1] Amorim D. de Souza 1994: A new
suprageneric classification of the Scatopsidae (Diptera:
Psychodomorpha). Iheringia, Série Zooogia 77: 107-112.
[2] Chandler P. 1998:
Scatopsidae. In Chandler P. (ed.): Checklists of Insects of the
British Isles (New Series). Part 1. Diptera (Incorporating a List of
Irish Diptera). Handbooks for the Identification of British
Insects, Vol. 12. Royal Entomological Society, London, pp.
47-48.
[3] Haenni J.-P. 1997:
Family Scatopsidae. In Papp L. & Darvas B. (eds): Contributions
to a Manual of Palaearctic Diptera (with special reference to flies
of economic importance). Volume 2. Nematocera and Lower Brachycera.
Science Herald, Budapest, pp. 255-272.
[4]
Haenni J.-P.
1998: Scatopsidae. In: Merz B., Bächli G., Haenni J.-P. & Gonseth Y. (eds):
Diptera
– Checklist. Fauna Helvetica 1.
CSCF/ SEG,
Neuchâtel, pp. 141-144.
[5] Haenni J.-P. 2002:
Redescription de Scatopse filamentosa Duda, 1928, avec une
nouvelle synonymie (Diptera, Scatopsidae). Bulletin de la Société
Neuchâteloise des Sciences Naturelles 125: 93-96.
[6] Haenni J.-P. 2006: Scatopsidae. In: Jedlička L., Stloukalová V.
& Kúdela M. (eds): Checklist of Diptera of the Czech
Republic and Slovakia. Electronic version
1. <http://zoology.fns.uniba.sk/diptera>. ISBN
80-969629-0-6. Retrieved 04.06.2009.
[7] Haenni J.-P. 2007: Scatopsidae. In Jong H. de (ed.): Fauna Europaea:
Diptera, Nematocera.
Fauna Europaea version 1.3.
<http://www.faunaeur.org>.
Retrieved 04.06.2009.
[8] Haenni J.-P.
2009: Scatopsidae. In Roháček J. & Ševčík J. (eds): Diptera of the Poľana
Protected Landscape Area – Biosphere Reserve (Central Slovakia). SNC SR,
Administration of the PLA – BR Poľana, Zvolen, pp. 94-96.
[9] Haenni J.-P. & Barták M. 2000:
Scatopsidae. In Barták M. & Vaňhara J. (eds): Diptera in an
industrially affected region (North-Western Bohemia, Bilina and
Duchcov environs), I. Folia Facultatis Scientiarum Naturalium
Universitatis Masarykianae Brunensis, Biologia 104:
105-109.
[10]
Haenni
J.-P. & Barták M. 2006:
Faunistic records from the Czech Republic (Diptera:
Scatopsidae). Entomofauna carpathica 18: 85.
[11] Haenni J.-P., Barták M. & Kubík Š.
2005: Scatopsidae. In Barták M. & Kubík Š. (eds): Diptera of
Podyjí National Park and its Environs. Česká zemědělská
univerzita, Praha, pp. 90-93.
[12] Haenni J.-P. & Vaillant F. 1994:
Description of dendrolimnobiontic larvae of Scatopsidae (Diptera)
with a review of our knowledge of the preimaginal stages of the
family. Mitteilungen der Schweizerischen Entomologischen
Gesellschaft 67:
43-59.
[13] Kozánek M. & Roller L. 1997: A
study of scatopsid (Diptera: Scatopsidae) communities in Slovakia.
Biologia, Bratislava 52: 637-646.
[14] Martinovský J. 1997a: Contribution
to the knowledge of Scatopsidae (Diptera) from the Czech and Slovak
Republics, with descriptions of two new species. Časopis Slezkého
Muzea v Opave (A) 45 (1996): 193-210.
[15] Martinovský J. 1997b: Scatopsidae.
In Chvála
M. (ed.): Check List of Diptera (Insecta) of the Czech and Slovak
Republics. Carolinum – Charles University Press, Prague, pp.
32-33.
[16] Martinovský J. 1998. Scatopsidae.
In Rozkožný R. & Vaňhara J. (eds): Diptera of the Pálava Biosphere
Reserve of UNESCO, I. Folia Facultatis Scientiarum Naturalium
Universitatis Masarykianae Brunensis, Biologia 99: 83-85. |
ASPISTINAE |
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Aspistes Meigen,
1818 |
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berolinensis
Meigen, 1818 |
CZ |
(B |
M |
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SK |
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ECTAETIINAE |
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Ectaetia Enderlein,
1912 |
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clavipes
(Loew, 1846) |
CZ |
(B |
M |
) |
SK |
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PSECTROSCIARINAE |
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Anapausis Enderlein,
1912 |
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albohalterata
Duda, 1928 |
CZ |
(B |
M |
) |
SK |
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dufourella
Haenni, 1984 |
CZ |
( |
M |
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SK |
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floricola
Chandler, 1999 |
CZ |
(B |
M |
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haemorrhoidalis
Duda, 1928 |
CZ |
(B |
M |
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SK |
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helvetica
Haenni, 1984 |
CZ |
(B |
M |
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SK |
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inermis
(Ruthé, 1831) |
CZ |
(B |
M |
) |
SK |
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nigripes
(Zetterstedt, 1860) |
CZ |
(B |
M |
) |
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rectinervis
Duda, 1928 |
CZ |
(B |
M |
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SK |
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rohaceki
Martinovský, 1997 |
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SK |
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soluta
(Loew, 1846) |
CZ |
(B |
M |
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SK |
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talpae
(Verrall, 1912) |
CZ |
(B |
M |
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SK |
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SCATOPSINAE |
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COLOBOSTEMATINI |
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in PCV2 all species in tribe SCATOPSINI [1] |
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Colobostema
Enderlein, 1926 |
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infumatum
(Haliday, 1833) |
CZ |
(B |
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nigripenne
(Meigen, 1830) |
CZ |
(B |
M |
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SK |
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obscuritarse
(Strobl, 1898) |
CZ |
( |
M |
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triste
(Zetterstedt, 1850) |
CZ |
(B |
M |
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SK |
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Efcookella Haenni,
1998 |
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albitarsis
(Zetterstedt, 1850) |
CZ |
(B |
M |
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SK |
in PCV2 in genus Cookella Freeman, 1985, preocc. [4] |
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Ferneiella Cook,
1974 |
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brevifurca
(Enderlein, 1912) |
CZ |
(B |
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incompleta
(Verrall, 1886) |
CZ |
(B |
M |
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SK |
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Holoplagia
Enderlein, 1912 |
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bullata
(Edwards, 1925) |
CZ |
(B |
M |
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SK |
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lucifuga
(Loew, 1870) |
CZ |
(B |
M |
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SK |
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richardsi
(Edwards, 1934) |
CZ |
( |
M |
) |
SK |
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transversalis
(Loew, 1846) |
CZ |
(B |
M |
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SK |
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RHEGMOCLEMATINI |
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in PCV2 as THRIPOMORPHINI, unnecessary change of name for RHEGMOCLEMATINI |
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Thripomorpha
Enderlein, 1905 |
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bifida
(Zilahy-Sebess, 1956) |
CZ |
(B |
M |
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SK |
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freyi
(Cook, 1969) |
CZ |
(B |
M |
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haennii
Martinovský, 1997 |
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SK |
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halteratum
(Meigen, 1838) |
CZ |
(B |
M |
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SK |
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paludicola
Enderlein, 1905 |
CZ |
(B |
M |
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SK |
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verralli
(Edwards, 1934) |
CZ |
(B |
M |
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SK |
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SCATOPSINI |
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Apiloscatopse Cook,
1974 |
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flavicollis
(Meigen, 1818) |
CZ |
(B |
M |
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SK |
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flavocincta
(Duda, 1928) |
CZ |
(B |
M |
) |
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gracilis
(Duda, 1928) |
CZ |
(B |
M |
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SK |
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filamentosa
(Duda, 1928) |
CZ |
( |
M |
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SK |
syn. hennigi Cook, 1974 [5] |
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mattheyi
Haenni, 1981 |
CZ |
( |
M |
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picea
(Meigen, 1818) |
CZ |
(B |
M |
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SK |
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scutellata
(Loew, 1846) |
CZ |
(B |
M |
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SK |
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styriaca
(Enderlein, 1926) |
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SK |
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Pharsoreichertella
Cook, 1974 |
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in PCV2 as subgenus in Reichertella Enderlein, 1912 [1] |
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hamifera
(Strobl, 1909) |
CZ |
(B |
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simplicinervis
(Duda, 1928) |
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SK |
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Reichertella
Enderlein, 1912 |
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in PCV2 all species in suppressed subgenus Reichertella Enderlein, 1912 [1] |
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geniculata
(Zetterstedt, 1850) |
CZ |
(B |
M |
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SK |
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nigra
(Meigen, 1804) |
CZ |
(B |
M |
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SK |
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pulicaria
(Loew, 1846) |
CZ |
(B |
M |
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SK |
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Scatopse Geoffroy,
1762 |
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lapponica
Duda, 1928 |
CZ |
(B |
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SK |
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notata
(Linné, 1758) |
CZ |
(B |
M |
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SK |
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SWAMMERDAMELLINI |
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Coboldia Melander,
1916 |
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fuscipes
(Meigen, 1830) |
CZ |
(B |
M |
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SK |
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Rhexoza Enderlein,
1936 |
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subnitens
(Verrall, 1886) |
CZ |
(B |
M |
) |
SK |
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Swammerdamella
Enderlein, 1913 |
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acuta
Cook, 1956 |
CZ |
(B |
M |
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SK |
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adercotris
Cook, 1972 |
CZ |
(B |
M |
) |
SK |
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brevicornis
(Meigen, 1830) |
CZ |
(B |
M |
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SK |
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genypodis
Cook, 1972 |
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SK |
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pediculata
(Duda, 1928) |
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SK |
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